Description:

Kadua formosa is a subshrub that can grow up to 1 meter tall, characterized by a prostrate main stem that is hollow and somewhat quadrangular or 4-costate, with a glabrous surface. The leaves are simple and opposite. The leaf blades are lanceolate to elliptic or oblong, measuring 5.5–14 cm in length and (1.6–)2.3–4.5 cm in width. The apex of the leaves is acuminate, and the base is cuneate. The leaves are coriaceous with entire margins, and the lateral veins are strongly ascending. The higher-order venation on the lower surface is conspicuously reticulate. Petioles are short, ranging from 0.2–0.4 cm in length. Stipules are broadly deltate, measuring 3.5–4.5 mm long, adnate to petioles, and forming a persistent sheath with a long acuminate apex. The flowers occur in dense corymbose inflorescences that are subtended by two cordate bracts, often with a lateral inflorescence in each. The central flower of the ultimate cyme is sessile or on a pedicel shorter than the two lateral cymes, with lateral pedicels approximately 1–2 mm long. The hypanthium is cylindrical-turbinate, about 2 mm long. Flowers are insect-pollinated, bisexual (perfect), sometimes functionally unisexual, and often heterostylous. Calyx lobes are bluntly deltate, small, and do not enlarge in fruit. The corolla is white, fleshy, and salverform with a tube 20–26 mm long, and the lobes are approximately 6–7 mm long. A nectary disc is present. Stamens are alternate with the corolla lobes and inserted somewhat below the corolla throat; anthers are sessile or on short filaments and dithecal, opening by longitudinal slits. The ovary is inferior or partly so, 2(-4)-locular with few to numerous ovules on fleshy placentas near the middle of the septum. The style is as many as carpels, glabrous, with lobes flattened, terminal, and slender; the stigma is bilobed or subcapitate, included or exserted. The fruits are ellipsoid–cylindrical capsules, measuring 9–12 mm long and 3–3.5 mm in diameter. They dehisce loculicidally across the disk and then septicidally with a heavily sclerified endocarp and a depressed–conical disk. Seeds are dark brown to nearly black, peltate, angled, and contain a well-developed oily endosperm, although the endosperm is occasionally scanty or absent. It is known only from wet cliffs on Maui.

Kadua formosa

Family: Rubiaceae

Hawaiian Name: N/A

 Minimum Length: 0.49 (mm)

 Maximum Length: 0.65 (mm)

 Minimum Width: 0.37 (mm)

 Maximum Width: 0.49 (mm)

 Minimum Mass: 0.014 (mg)

 Maximum Mass: 0.030 (mg)

 Texture: TBD

Geographic Distribution:

 Maui
Species Of Concern

PEPP Status:

Not Applicable

Native Status:

Endemic

2D Seed Shapes

Bean-shaped (Reniform): Resembling the shape of a kidney or bean.

Square or Oblong: Angular shape with all sides more or less equal or longer than wide.

Triangular: Three-sided shape, broadest below the middle.

Elliptic: Rounded at both ends. shaped like an ellipse or oval.

Ovate or Obovate: egg-shaped, broader at one end.

Linear: Extremely long and narrow, with sides usually parallel.

Round: Appears circular, with length and width roughly the same.

Cardioid (Heart-Shaped): One edge has a prominent double curve into two lobes.

3D Seed Shapes

Cylindrical: A cylinder with parallel sides and a circular cross-section.

Discoid: Resembling a disc or plate with two relatively flat surfaces.

Fusiform: Spindle-shaped, tapering at both ends.

Globose: Spherical.

Lenticular: Resembling a lens with two convex surfaces.

Ovoid: Egg-shaped.

Plano-convex: Having a flat ventral (planar) surface and a curved dorsal (convex) surface.

Polygonal: Having more than three flat faces meeting at angles.

Sectoroid: Resembling a segment of an orange with two relatively flat faces and one curved face.

Subglobose/ellipsoid: Inflated shape but less than spherical.

Trigonous: Having three faces that meet at a distinct angle; triangular in cross-section.

Federal Status

Endangered species (E) are species that are so rare they’re in danger of becoming extinct.

Threatened species (T) are species that are likely to become endangered within the foreseeable future throughout all or a significant portion of their range.

Species of Concern (SOC) are species about which there are some concerns regarding status and threats, but insufficient information is available to list the species under the Endangered Species Act (ESA).

Plant Extinction Prevention (PEP) is a state status for rare plants that focuses its efforts on species that have fewer than 50 remaining wild individuals.

Storage Behavior

Orthodox seeds (desiccation-tolerant): can be dried to an internal seed moisture of less than 12% water and stored at freezing temperatures.

Intermediate seeds (temperature-sensitive) tend to age faster than orthodox seeds and may have only a 5-year lifespan when stored at -20°C. They are recommended for storage at 5°C.

Recalcitrant seeds (desiccation-sensitive): cannot be stored in a conventional freezer as they cannot survive after drying and/or freezing at -20°C. They are recommended for cryopreservation storage.